1 Introduction 
The first Greeks to inhabit Aetolia were  those who migrated there from Elis, led by Aetolus, son of Endymion. When they  migrated, Aetolia was inhabited by the Curetes. [1] 
Strabo reports that some Hyantes, chased by  Cadmus, migrated from Boeotia to Aetolia. [2] 
Curetes is presumed to have been Hyantes with  a name change. 
The marriage of Pleuron, son of Aetolus, to a  Curetes woman was the cause of a long conflict in Aetolia. 
      2 Age of Aetolus, son of Endymion 
        2.1 Emigration from Elis 
        In 1320 BC, Aetolus, son of Endymion of Elis in  Eleia, led the Epeans across the sea north from the Peloponnesus peninsula. [3] 
        Aetolus was chased from Elis by Salmoneus of  Salmone. [4] 
        Aetolus was king of Elis, but his successor  was Eleius, son of his sister Eurycyda. [5] 
        It is assumed that Eleius' father was Alector  (or Alexinus), son of Salmoneus. [6] 
        So, it seems that there was a power struggle  in Elis, and Salmoneus, who wanted to put his grandson on the throne, expelled  Aetolus and his supporters from Elis. 
      2.2 Founding of Pleuron 
        Aetolus settled at the foot of Mount Curium,  between the rivers Achelous and Euenus, and founded Pleuron. [7] 
        The land was called the Curetian country, and  the indigenous people were the Curetes. [8] 
        Aetolus fought against the Curetes and won,  but was unable to drive them far. [9] 
        The region Aetolus settled in was named after  him, Aetolia, and the people who lived there became known as Aetolians. [10] 
      2.3 Wife of Aetolus 
        Before Aetolus emigrated, he had married  Pronoe, daughter of Phorbas, and they had two sons, Pleuron and Calydon. [11] 
        Pronoe, the wife of Aetolus, was the daughter  of Phorbas, son of Lapithus (or Lapithes). [12] 
        Lapithus was the progenitor of the Lapiths  and the son of Aeolus, the son of Hippotes, the son of Mimas. 
        Aetolus was the son of Endymion, the son of  Aethlius, the son of Aeolus, the son of Hippotes, the son of Mimas. 
        In other words, Aetolus and Pronoe were second  cousins. 
      3 Age of Pleuron, son of Aetolus 
        3.1 Marriage of Pleuron 
        Pleuron married Xanthippe (or Demonice),  daughter of Dorus the Curetes, and succeeded her father, and lived in Pleuron.  [13] 
        Xanthippe appears to have been a prisoner of  war, which was common in this era, but looking at the relationship between the  descendants of Aetolus and Curetes, it seems that the marriage was for peace  between the two parties. 
      3.2 Founding of Calydon 
        Pleuron's brother Calydon moved about 12 km  east-southeast of Pleuron and founded Calydon. 
        The Epeans (Aeolis), who migrated from Elis  with Aetolus, lived in Pleuron, adjacent to the settlement of Curetes. However,  they were gradually oppressed by the Curetes and moved to Calydon. [14] 
      3.3 Marriage from Eleia 
        In 1301 BC, the alarmed Calydon took as his  wife Aeolia, the daughter of Amythaon of Pylus, who became ruler of Eleia in  place of Salmoneus, who had driven his father Aetolus from Elis. [15] 
        It is assumed that many Aeolis migrated from  Elis to Calydon, accompanying Aeolia. 
        Aeolia was the daughter of Amythaon, the son  of Tyro, the daughter of Salmoneus, the brother of Aethlius, the father of  Endymion, the father of Aetolus, the father of Calydon. So, Aeolia was  Calydon's third cousin. 
      4 Age of Thestius, son of Pleuron 
        4.1 Father of Diomedes' grandfather Oeneus 
        4.1.1 The legendary father of Oeneus 
        According to tradition, the father of Oeneus  is said to be: 
        Homer is Portheus, father of Agrius and  Melas. [16] 
        Hyginus is Parthaon. [17] 
        Apollodoros is Parthaon, son of Agenor, son  of Pleuron. [18] 
        Antoninus Liberalis is Portheus. [19] 
        Pausanias is Porthaon. [20] 
      4.1.2 Another Oeneus 
        Pausanias records the tradition of Mothone of  Messenia, that when Oeneus returned to Diomedes after the capture of Troy, his  illegitimate daughter Mothone was born. [21] 
        However, Oeneus' grandson Diomedes was  married before going to Troy, and Oeneus after the Trojan War is old enough to  have his great-grandson. It is hard to believe that a daughter was born to the  elderly Oeneus. [22] 
        Additionally, Homer writes in the Catalogue  of Ships that Oeneus is already dead. [23] 
        Taking these traditions together, it seems  true that Parthaon, son of Agenor, son of Pleuron, had a son named Oeneus, but  he seems to be a different person from Diomedes' grandfather. 
      4.1.3 Estimated father of Oeneus 
        According to the Mothone tradition,  Parthaon's son Oeneus was a contemporary of Diomedes, and Diomedes' grandfather  Oeneus was a contemporary of Parthaon's son Oeneus' grandfather Agenor. 
        Diomedes' grandfather Oeneus is therefore  presumed to have been the son of Pleuron or of Pleuron's brother Calydon. 
        It is also said that a quarrel arose between  the Curetes and the Calydonians, and Meleager, the son of Oeneus, killed the  sons of Thestius, the son of Pleuron. [24] 
        This means that Oeneus, the ruler of Calydon,  is not of Curetes blood. 
        Therefore, the father of Diomedes'  grandfather Oeneus was not Pleuron, son of Aetolus, but Calydon, son of  Aetolus. 
        Pleuron, son of Aetolus, married Xanthippe,  daughter of Dorus the Curetes, and his sons Agenor and Thestius were Curetes. [25] 
      4.1.4 Marriage of Oeneus 
        In 1267 BC, Oeneus took Althaea, daughter of  Thestius of Pleuron, as his wife. [26] 
      4.2 Father of Thestius 
        As for Thestius, the father of Leda, the wife  of Tyndareus, son of Oebalus, Pausanias alone tells us that he was Agenor, son  of Pleuron. [27] 
        However, Agenor seems not to be Thestius'  father for the following reasons. 
        If Thestius' father is Agenor, Agenor's son  Porthaon becomes Thestius' brother. [28] 
        In the story of the Calydonian Boar Hunt,  nine of Thestius's eleven sons are named, but Porthaon's five sons are not. 
        The story mentions the sons of Oeneus, son of  Calydon, son of Aetolus, so Thestius was a contemporary of Oeneus. 
        Thestius was the ruler of Pleuron who was at  war with Calydon, and it is assumed that Thestius' father was Calydon's brother  Pleuron. [29] 
      4.3 Marriage to Ozolian Locris 
        In 1266 BC, Calydon's daughter Protogenia  married Andraemon, son of Aetolus, who lived in Amphissa. [30] 
        It is assumed that Calydon tried to recruit  the neighboring Ozolian Locrians to his side in order to counter Pleuron. 
      4.4 Founding of Olenus 
        In 1265 BC, Hipponous, son of Dexamenus,  migrated from Olenus in Achaia to Aetolia and founded Olenus. [31] 
        Pleuron, near that town, was the birthplace  of Molione, the wife of Actor, the brother of Dexamenus, the father of  Hipponous. [32] 
      4.5 Emigration from Sparta 
        In 1265 BC, Tyndareus, son of Oebalus,  migrated from Sparta to Pleuron. [33] 
        Thestius of Pleuron where Tyndareus migrated, was the husband of  Aphareus' sister Demonice. [34] 
        Demonice was Tyndareus' half-sister. [35] 
      4.6 Emigration from Messenia 
        Tyndareus went to Arene in Messenia before  going to Aetolia. Idas, the son of Aphareus, who lived in Arene, was a  contemporary of Tyndareus, and the two emigrated together. [36] 
        Idas' mother Arene was Tyndareus' sister, and  Idas was Tyndareus' nephew. 
        Idas married Marpessa, daughter of Evenus of  Pleuron. [37] 
        According to legend, Idas took Marpessa and  was chased by Evenus, but Evenus could not catch Idas and threw himself into  the river, which became known as Evenus. [38] 
        The Evenus River flows from Pleuron beyond  Calydon, and Idas may have fled from Pleuron to Calydon. 
        Later, the daughter of Idas married Meleager,  son of Oeneus of Calydon. [39] 
      4.7 Emigration from Argos 
        The Hyettus affair at Argos led to a conflict  between Melampus' two sons, Abas and Mantius. [40] 
        In 1264 BC, Mantius, son of Melampus, was  chased from Argos by Abas and fled to Calydon, the marriage of his aunt Aeolia.  [41] 
        Oecleus, son of Mantius, married  Hypermnestra, daughter of Thestius of Pleuron, and they had a son, Amphiaraus.  [42] 
      5 Age of Oeneus, son of Calydon 
        5.1 Marriage of Idas' daughter 
        In 1247 BC, Cleopatra (or Halcyone), daughter  of Idas, married Meleager, son of Oeneus of Calydon. [43] 
      5.2 Emigration to Argos 
        In 1247 BC, Mantius, son of Melampus,  returned from Calydon to Argos with his son Oecles and his grandson Amphiaraus.  [44] 
        After this, a battle begins in Aetolia  between Calydon and Pleuron. 
        The disappearance of Mantius' son Oecles and  his wife Hypermnestra, who were related to the two towns, seems to have  reignited the conflict between the two towns. 
        It is also said that Marpessa, Idas' wife  from Pleuron, committed suicide, which may have something to do with the cause  of the battle. [45] 
      5.3 Battle of Calydon and Pleuron 
        The conflict between settlers and indigenous  peoples that began in 1246 BC, during the time of Aetolus's settlement, became  a battle that has been passed down to future generations. The fierce battle  between the Aetolians and the Curetes lasted for six days. [46] 
      5.3.1 Story of the Calydonian boar hunt 
        The story of the Calydonian boar hunt  features many of the heroes who took part in the Argonauts' expeditions. [47] 
        Bacchylides, one of the nine poets of ancient  Greece, also tells of the battle between the Curetes and the Aetolians.  However, heroes do not appear in his poems. In the 5th century BC, the story is  thought to have survived before many heroes were added. [48] 
      5.3.2 Participants in the battle 
        Tyndareus, son of Oebalus, and Idas, son of  Aphareus, also took part in this battle. 
        Tyndareus was on the side of Pleuron, Idas on  the side of Calydon. Later, Tyndareus and Idas moved the battle to  Peloponnesus, and it became a battle between Messenia and Laconia. [49] 
      5.3.3 Destruction of Olenus 
        Oeneus of Calydon attacked Olenus of Aetolia,  captured it, and took Periboea, daughter of Hipponous, prisoner. [50] 
        Strabo notes that Aeolis was the one who  destroyed Olenus. [51] 
        Oeneus belongs to the Aetolians, a branch of  the Epeans, a branch of the Aeolis, and the Aeolis that Strabo mentions are  also like Oeneus. 
        However, the name of Olenus of Aetolia  appears in Homer's Catalogue of Ships, so Oeneus is not the destroyer. [52] 
        The destroyer of Olenus is presumably Aeolis,  who lived in Calydon and Pleuron in the 5th century BC, as described by  Thucydides. [53] 
      5.3.4 Victory or defeat in battle 
        Oeneus of Calydon was victorious and took  control of Pleuron and Curetes. 
        Eight years later, Heracles pleased Oeneus by  reclaiming Paracheloitis near the mouth of the Achelous River. [54] 
        Oeneus' rule at this time extended westward  from Calydon, far beyond the settlements of Pleuron and Curetes, to the river  Achelous, which bordered Acarnania. [55] 
        The name of the river Achelous came after the  time of Alcmaeon, son of Amphiaraus, and in the time of Heracles it was called  the river Thestius. [56] 
      5.3.5  Inhabitants of Pleuron 
        The  descendants of Thestius, defeated in the battle, migrated inland from Pleuron  and founded Thestia. [56-1] 
        Pleuron  continued to be inhabited by Parthaon, son of Agenor, son of Pleuron. 
      5.4 Emigration to Messenia 
        In 1245 BC, Aphareus' son Idas returned from  Calydon to Arene in Messenia and succeeded his father. [57] 
      5.5 Emigration from Arcadia 
        In 1238 BC, Heracles, son of Amphitryon,  migrated from Pheneus in Arcadia to Calydon. [58] 
        Shortly before this, Heracles had been at war  with Elis, and Oecles, son of Mantius, who led the Argives, had also taken part  in the battle. [59] 
        It is assumed that Heracles was introduced to  Oeneus of Calydon by Oecles. 
        Oecles' wife Hypermnestra was the daughter of  Thestius, and Oeneus was the husband of her sister Althaea. [60] 
        In other words, Oeneus was Oecles'  brother-in-law. 
        Melampus, the father of Mantius, the father  of Oecles, was also the brother of Aeolia, the wife of Calydon, the father of  Oeneus. [61] 
        In other words, Oeneus was also the son of  Oecles' grandfather's brother-in-law. 
      5.6 Expedition to Thesprotia 
        In 1237 BC, the Calydonians led an expedition  into Thesprotia and captured Ephyra, led by Heracles. [62] 
        There was no conflict of interest between  Calydon and the distant Ephyra, and the Calydonians participated in Heracles'  expeditions. 
      5.7 Emigration to Sparta 
        In 1237 BC, Tyndareus migrated from Aetolia  to Sparta. Legend has it that Heracles recalled Tyndareus to Sparta after his  battle with Hippocoon. [63] 
        However, Tyndareus returned to Sparta because  Hippocoon and Icarius were gone, and there was no one to succeed Oebalus. 
        Tyndareus' two sons, Castor and Polydeuces,  who were born and raised in Aetolia, also migrated to Sparta. Many legends tell  of Dioscuri joining the expeditions of the Argonauts from Sparta. However, if  Dioscuri really joined the expedition, it was not from Sparta, but from  Pleuron. 
      5.8 Emigration to Trachis 
        In 1235 BC, Heracles migrated from Calydon to  Trachis. [64] 
        Legend has it that Heracles decided to leave  Calydon after he accidentally killed the child of a relative of Oeneus. [65] 
        However, this relocation of Heracles seems to  have been influenced by the will of Eurystheus of Mycenae, who feared that  Heracles would further expand his influence in Calydon. It is presumed that it  was Amphiaraus, son of Oecles, who was born in Calydon and lived in Argos, who  conveyed Eurystheus' intentions to Oeneus. Oeneus and Oecles were stepbrothers  through their respective wives. [66] 
      5.9 Exile to Argos 
        In 1226 BC, Tydeus, son of Oeneus, went into  exile to Adrastus, son of Talaus, in Argos. [67] 
        Oeneus' mother Aeolia was the sister of Bias,  father of Talaus, father of Adrastus, and Tydeus was Adrastus' second cousin. 
        Tydeus married Deipyla, daughter of Adrastus,  and they had a son, Diomedes. [68] 
      5.10 Marriage to Amphissa 
        In 1222 BC, Gorges (or Gorge), daughter of  Oeneus of Calydon, married Andraemon (or Andrawmon) of Amphissa in Ozolian  Locris. [69] 
        The husband of Oeneus' sister Protogenia was  Andraemon, son of Aetolus, who lived in Amphissa. [70] 
        Andraemon and Gorges had a son, Thoas. [71] 
      5.11 Exile to Argos 
        In 1202 BC, Oeneus was driven from Calydon by  the sons of Agrius and exiled to Diomedes of Argos. [72] 
        Oeneus' son-in-law Heracles had already died,  but in Amphissa was his son-in-law Andraemon. At that time, the Argives had  captured Thebes, and the fame of Argos seems to have reached Oeneus. Oeneus was  in exile, his son Tydeus already dead and relying on his grandson Diomedes. 
      5.12 Diomedes' Expedition 
        In 1202 BC, Diomedes, son of Tydeus, son of  Oeneus, went on an expedition to Aetolia to avenge Oeneus's exile from Calydon.  [73] 
      5.12.1 Expedition Participants 
        Different traditions say that Diomedes'  companions in the expedition were either Alcmaeon, son of Amphiaraus, or  Sthenelus, son of Capaneus. [74] 
        Alcmaeon's mother, Hypermnestra, was the  daughter of Thestius of Pleuron, and it is unlikely that Alcmaeon participated  in battles against his kin. [75] 
        Sthenelus was a close friend of Diomedes, and  it is believed that Sthenelus assisted Diomedes in his expeditions. [76] 
        Also participating in the expedition was  Andraemon of Amphissa in Ozolian Locris. 
        Andraemon was the husband of Gorges, the  daughter of Oeneus, and the brother-in-law of Tydeus, the father of Diomedes.  [77] 
        After the battle, Diomedes left Aetolia in  charge of Andraemon. [78] 
        There is also a legend that Oeneus  accompanied him on this expedition and returned to Argos again. [79] 
        However, if that were the case, the  expedition to bring Oeneus home would be meaningless. Oeneus had died before  this, and Diomedes is presumed to have gone on an expedition to avenge Oeneus. 
      5.12.2 Descendants of Agrius  
        The descendants of Agrius, defeated  by Diomedes, migrated inland from Pleuron and founded Agrinium. [80] 
        Agrinium belonged to the  Eurytanians, who had the largest population among the Aetolians. [81] 
        Thucydides notes that the  Eurytanians were the most uncivilized of the Aetolians, speaking a difficult  language and eating raw meat. [82] 
        Agrinium was still in existence  when Philip, king of Macedonia, attacked the Aetolians in 218 BC. [83] 
      5.13 Marriage to Acarnania 
        In 1202 BC, Callirhoe, daughter of Achelous  and Melpomene (or Sterope), married from Pleuron to Alcmaeon who lived in Argos-Amphilochicum  in Acarnania. [84] 
        Hypermnestra, the wife of Oecles, the father  of Amphiaraus, the father of Alcmaeon, was a cousin of Parthaon, the father of  Melpomene, the mother of Callirhoe. 
      6 Age of Thoas, son of Andraemon 
        6.1 Trojan War 
        Andraemon and Gorge's son Thoas led the  Aetolians in the expedition to Troy. [85] 
        It is likely that Thoas' force also included  the people of Ozolian Locris, including Amphissa, but Homer does not mention  Amphissa. 
        It is assumed that Thoas migrated from  Amphissa to Calydon, taking many Aeolis with him to rule the Aetolians. [86] 
        It is believed that Amphissa had a small  population in the Trojan War Era, and it is assumed that the original author of  the Iliad was aware of the actual situation at that time. 
      6.2 Emigration to Italy 
        In 1184 BC, Diomedes, son of Tydeus, migrated  from Argos through Aetolia to the eastern coast of the Italian peninsula and  founded Argyrippe. [87] 
        Diomedes married Euippe, daughter of Daunius  (or Daunus), king of the Daunians, and they had two sons, Diomedes and  Amphinomus. [88] 
        Diomedes also founded Arpi and Beneventum.  [89] 
        The Aetolians, who migrated with Diomedes,  founded Brundisium in the southeastern part of the Italian peninsula. [90] 
      7 Age of Haemon, son of Thoas 
        7.1 Emigration from Italy 
        After the death of Diomedes in 1150 BC, his  son Amphinomus emigrated from the Italian peninsula to Aetolia. [91] 
        It is assumed that Haemon, son of Thoas,  accepted Amphinomus and married his son to Amphinomus' daughter Thyrie. 
        Cycnus, son of Thyrie, lived near Pleuron.  [92] 
      7.2 Emigration to Argos 
        In 1120 BC, Erginus, possibly the son of  Amphinomus, moved to Argos, the hometown of his grandfather Diomedes. [93] 
        In 1110 BC, Erginus aided Temenus by stealing  the statue of Palladium, the city's patron god, from Argos during the  Heracleidae's siege of the city. [94] 
        Erginus and Temenus were related through  their common ancestor Oeneus of Calydon. 
      7.3 Emigration to Elis 
        In 1105 BC, Oxylus, son of Haemon, son of  Thoas, led the Aetolians in migrating from Aetolia to Elis. [95] 
        Those who remained in Calydon and Pleuron  were the Aeolis, who had migrated from Amphissa, and the Aetolians, who had  migrated from the Italian peninsula with Diomedes's son Amphinomus after the  death of Diomedes. 
      8 Age after Heracleidae's return 
        Afterwards a conflict arose between the  Aeolis and the Aetolians, and the descendants of Diomedes migrated to the west  side of the Achelous River, to Acarnania. 
        Acarnania was inhabited by people who had  emigrated from Thebes after being defeated by Epigoni, creating Astacus. Also,  people who migrated from Argos founded and lived in Argos-Amphilochicum near  the Ambracian Bay. [96] 
        The descendants of Diomedes called themselves  Oeniadae and became part of the Acarnanians rather than the Aetolians. [97] 
        The inhabitants of Calydon and Pleuron called  themselves Aeolis to distinguish themselves from other Aetolians. [98] 
      9 Peloponnesian War Era 
        9.1 Offensive and defensive battle of  Oeniadae 
        In 459 BC, during a war with the Spartans,  the Messenians, holed up on Mount Ithome, were chased by the Lacedaemonians and  moved to Naupactus in the Ozolian Locris. [99] 
        Naupactus was acquired by the Athenians, who  drove out the Ozolian Locrians, due to their hatred of the Spartans, and  offered it to the Messenians. [100] 
        The Messenians of Naupactus, seeking to repay  the Athenians' favor, laid siege to the Oeniadae, which were hostile to the  Athenians, and, after a truce, evacuated the inhabitants. [101] 
        Driven from their homes, the Oeniadae, with  the help of other Acarnanians, laid siege to the town and recaptured it eight  months later. 
        In 455 BC, Pericles of Athens recruited the  Acarnanians, except for the Oeniadae, and the following year he besieged  Oeniadae with the Aetolians, but was unable to capture it. [102] 
        In 424 BC, the Athenians formed the  Acarnanian League with the addition of the Oeniadae. [103] 
      9.2 Inhabitants of Calydon 
        In 405 BC, the Spartans defeated the  Athenians at the Battle of Aegospotami and expelled the Messenians from  Naupactus. [104] 
        In 389 BC, the Achaeans of Calydon, oppressed  by the Athenians allied with the Arcarnanians and Boeotians, reportedly  requested the aid of the Spartans. [105] 
        It is likely that the Spartans who expelled  the Messenians from Naupactus also expelled the inhabitants of Calydon and gave  it to the Achaeans. [106] 
        It is assumed that Andraemon, or Aeolis who  migrated from Amphissa to Aetolia with Thoas, son of Andraemon, was expelled by  the Spartans in 405 BC. 
        However, it appears that many of their  residents continued to live there. 
        In 29 BC, when the Roman emperor Augustus  tried to evict the Aetolians from their settlements and relocate them to the  new city in preparation for the construction of Nicopolis, most of the  inhabitants fled to Amphissa. [107] 
      10 Subsequent Oeniadae 
        In 331 BC, Oeniadae was attacked by the  Aetolians, who destroyed the town and expelled its inhabitants. [108] 
        These Aetolians are presumed to be Curetes  who live on the east side of the Achelous River, that is, in Aetolia. Among the  names of Aetolians tribes of the Roman period enumerated by Strabon is Curetes.  [109] 
        This attack by the Curetes appears to have  been in response to the rebellion against Macedonia led by Agis III of Sparta.  [110] 
        In 324 BC, Alexander the Great issued the decree for the recall of the exiles, which was  strongly opposed by the Aetolians, who had expelled the Oeniadae and made them  their own territory. Together with the Athenians, who distributed Samos among  the citizens, they became the spearhead of the anti-Macedonian movement. [111] 
        The name Oeniadae seems to have remained in  the Roman period, but details about its inhabitants are unknown. [112] 
        The Battle of Curetes, which began with the  settlement of Aetolus of Elis in 1320 BC, lasted for nearly 1,000 years. In the  end, it was a victory for the Curetes, who continued to live on the same land  until the end. 
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